Kaloula laosensis Poyarkov, Orlov, Gorin & Milto 2024, sp. nov.
Year: 2024
Type: other
Abstract: Kaloula laosensis Poyarkov, Orlov, Gorin & Milto sp. nov. Figs 2, 9 A, 10, 11, 12, Table 4 Chresonymy. Kaloula baleata [partim] — Orlov and Ananjeva (2007: 148); Nguyen et al. (2009: 94). Kaloula indochinensis [partim] — Chan et al. (2013: 334, 2014: 577); Teynié et al. (2014: 29); Chandramouli and Prasad (2018: 52); Poyarkov et al. (2021 b: 39); Holden (2023: 148). Holotype. ZISP 15284 (field label ZISP 199), adult male, Na Home Village, Bouphala District, near the border of Nakai-Nam Theun National Park, Khammouane Province, Laos, collected by N. L. Orlov, S. N. Nguyen, and K. D. Milto on June 18, 2009 (17.544528 ° N, 105.695278 ° E; elevation 174 m a. s. l.). Paratype (n = 1). ZMMU A-8144 (field label ZISP 200), adult female from the same location and with the same collection data as the holotype. Diagnosis. Kaloula laosensis sp. nov. is distinguished from its congeners by the following combination of morphological attributes: (1) medium size (SVL ranging 44.0– 54.8 mm); (2) eyes comparatively large (eye length comprising 75 % – 100 % of snout length); (3) dark-brown dorsally, with numerous irregular dark blotches forming pericloacal ring; (4) gray or beige ventrally with dense white mottling on belly and limbs; (5) orange triangular patch on either side of the neck posterior to eyes; (6) bright-orange axillary patch present; (7) orange inguinal patch present; (8) bright-orange butterfly-shaped blotch above cloaca present; (9) gray spot on tibiotarsal articulation present; (10) dark interorbital bar present; (11) enlarged, widened finger disks (7.8 % – 7.9 % of SVL), ca. 1.53 times wider than toe disks; (12) finger subarticular tubercle formula: 1: 1: 2: 2; (13) toe subarticular tubercle formula: 1: 1: 2: 2: 2; (14) three metacarpal tubercles, median metacarpal tubercle in contact with inner metacarpal tubercle; (15) two metatarsal tubercles, inner metatarsal tubercle ca. three times larger than outer metatarsal tubercle; (16) three small supernumerary tubercles at the basis of toes I, III, IV. Description of the holotype (Fig. 10). Adult male in a good state of preservation, habitus robust; head wider than long (HW / HL 1.19); snout projecting beyond lower jaw, truncated in dorsal and lateral views (Fig. 10 C); top of head flat; upper eyelid lacking supraciliary tubercles; eye length less than snout length (EL / SL 0.75) and less than interorbital distance (EL / IOD 0.77); pupils round; nostrils rounded, placed more towards the lateral sides of snout, located closer to tip of snout than to eye, relatively close to each other (IND / IOD 0.63); supratympanic fold flat, tubercular; tympanum not visible (Fig. 10 C); dorsal surfaces of body and limbs with sparse tubercules, getting denser backwards; ventral surfaces of body and limbs almost smooth (Fig. 10 A, B). Cloacal opening unmodified, directed posteriorly. Forelimbs relatively long, more than a half of hind limb length (FLL / HLL 0.63); hand long, comprising more than a half of lower arm length (HAL / LAL 0.6) and almost half of forelimb length (HAL / FLL 0.47); fingers rather robust, flattened in cross section; relative finger lengths: I <II <IV <III; no webbing between fingers; terminal digits flattened into very wide transversely expanded T-shaped disks (Fig. 10 E); finger subarticular tubercles distinct, protuberant, large and round, finger subarticular formula: 1: 1: 2: 2; metacarpal tubercles three (Figs 9 A, 10 E), inner metacarpal tubercle elongate, flattened; outer metacarpal tubercule oval, dilated, slightly bigger than inner (OPTL / IPTL 1.05), median metacarpal tubercle the smallest, oval-shaped, contacting both the inner and outer metacarpal tubercles (Figs 9 A, 10 E); hindlimbs robust, relatively short, not much longer than body length (HLL / SVL 1.12); relative toe lengths: I <II <V <III
Authors: Gorin, Vladislav A., Orlov, Nikolai L., Bragin, Andrey M., Pawangkhanant, Parinya, Milto, Konstantin D., Le, Dac Xuan, Nguyen, Tan Van, Dufresnes, Christophe, Suwannapoom, Chatmongkon, Poyarkov, Nikolay A.
Language: English
Subfield: Global and Planetary Change
Field: Environmental Science
Domain: Physical Sciences
Open Access status: green
OpenAlex ID: W6930794081
Cites:
Cited by:
Related to: