Reductodrilus maculatus Packova, Vorst & Kundrata 2026, sp. nov.
Year: 2026
Type: article
Abstract: Reductodrilus maculatus Packova, Vorst & Kundrata sp. nov. Figs 4, 5, 6 Ototretadrilus philippinus ab. maculata Wittmer, 1941: 197 [unavailable name, ICZN 1999: Art. 45.6. 2; see Keller and Hinson 2023]. Type material. Holotype: male, “ Boettcher, Phil.: Socorro, [Bucas Grande] 19.X.1916 // HOLOTYPE: Ototetradrilus [sic!] philippinus 1939, W. Wittmer, ab. maculatus Wittm. // O. philippinus ab. maculatus Wittm., det. W. Wittmer // Ototretadrilus philippinus maculatua [sic!] Wittmer, 1941, ZMAN type COLE.1031.1 // RMNH.INS.1092883 [with a barcode] ” (RMNH). Type locality. Philippines: Socorro. Diagnosis and comparison. This species can be recognized based on the following unique combination of characters: dorsum dark brown except for the lateral portions of pronotum and outer basal portions of elytra (Fig. 4 A), antennomeres 4–10 ~ 1.7–1.8 × as wide as long (Fig. 5 C), pronotum ~ 1.45 × wider than long (Fig. 5 E), elytra ~ 1.4 × as long as wide at humeri and 2.2 × as long as pronotum (Fig. 4 A), elytral pubescence bicolorous, with major portions of elytra having distinctly darker setae, and the aedeagal median lobe ~ 1.5 × as long as phallobase, not surpassing apices of parameres, widest basally, subparallel-sided for most of its length and then narrowed at apical 3 / 5 of their length (Fig. 6 D, F). While R. maculatus sp. nov. shares many external morphological characters with R. testaceiceps, it differs in much darker coloration of dorsum (Figs 7 A, 8 A), more strongly pectinate antennomeres 4–10 (1.7–1.8 × as wide as long versus 1.5–1.6 × in R. testaceiceps; Fig. 9 D), less transverse pronotum (~ 1.45 × wider than long versus 1.55–1.60 × in R. testaceiceps; Fig. 9 E), and different shape of male genitalia, with having relatively shorter median lobe and parameres compared to the phallobase, the median lobe widest basally (versus medially in R. testaceiceps; Fig. 10 D, F) and not surpassing apices of parameres. Redescription. Male. Body (Fig. 4 A, B) 7.1 mm long from frons to apex of abdomen and 5.0 mm long from frons to apex of elytra, 2.4 mm wide at humeri; pale to dark brown, head (except frontal part), antennae, palpi, pronotum medially from anterior to posterior margin, scutellum, elytra (except outer basal portions), major portions of tibiae (except basally), tarsomeres, and several last abdominal segments distinctly darker, dark brown; body surface covered with yellowish to dark brown setae (coloration of setae roughly corresponding to coloration of cuticle). Head (Fig. 5 A, B, D) including eyes ~ 0.7 × as wide as pronotum, dorsomedially very slightly convex, generally smooth, sparsely covered with fine punctures and short semi-erect setae mostly oriented anteriorly. Frontoclypeal region frontally abruptly declivous, antennal fossae separated by 0.8 × longitudinal diameter of fossa; rim dorsally forming narrow supra-antennal carina. Eyes moderately large, basally slightly emarginate. Labrum beneath (i. e., not on the same plane with frontoclypeal region), weakly sclerotized, sparsely covered with long suberect to erected setae. Mandibles unidentate, moderately long and robust, sickle-shaped, strongly curved; basally covered with sparse long setae. Maxillary palpi relatively short, 4 - segmented, basal palpomere shorter than wide, palpomere 2 slightly longer than wide; palpomere 3 shortest, wider than long; ultimate palpomere longest, apically dorsoventrally flattened and obliquely cut. Labial palpi minute, 3 - segmented; palpomere 2 short, transverse; ultimate palpomere longer than wide, similar in shape to ultimate maxillary palpomere. Antenna (Fig. 5 C) with 11 antennomeres, moderately long, slightly surpassing midlength of elytra (when positioned backwards), rather robust, scape longer than wide, antennomere 2 shortest, slightly wider than long; antennomeres 3–10 pectinate, antennomere 3 ~ 2.3 × as long as antennomere 2, ~ 1.5 × as long as wide, antennomeres 4–10 subequal in shape and length, ~ 1.7–1.8 × as wide as long; antennomere 11 ~ 2 × as long as antennomere 10, elongate, widest after half, apically rounded. Pronotum (Fig. 5 E) ~ 1.45 × wider than long, slightly convex, widest medially. Anterior margin widely rounded, with the edge slightly turned upwards; lateral margins slightly rounded; posterior margin slightly bisinuate, medially very slightly produced and shallowly emarginate. Anterior angles inconspicuous; posterior angles very short, almost rectangular, apically rounded. Surface of disc uneven, with distinct median longitudinal groove, one small posteromedial depression, and two large shallow anterolateral depressions, rather densely covered with punctures and setae; punctures moderately large, shallow, intervals between punctures usually ~ 1 puncture diameter, interstices rather smooth; setae moderately long, semi-erect, mostly oriented posteriorly, those near posterior margin oriented anteriorly. Lateral pronotal carina complete, visible in dorsal view, anteriorly continuous with anterior margin of pronotum. Hypomeron with surface mostly smooth and shiny, with shallow punctures and semi-erect setae. Prosternum distinctly transverse, with prosternal process reduced to short median protrusion. Scutellar shield (Fig. 5 E) tongue-shaped, rather large, almost as long as wide, widest anteriorly, gradually slightly narrowed toward apex, apex subtruncate; surface densely and coarsely punctate, punctures approx. the same size as those on elytra, and with short semi-erect setae. Mesoventrite short, roughly triangular. Metaventrite with distinct long discrimen; metepisternum distinctly wider basally and gradually narrowed apicad. Elytra (Fig. 4 A, B) both combined ~ 1.4 × as long as wide, 2.2 × as long as pronotum, strongly dehiscent, not meeting along suture, sides slightly sinuate, apices separately rounded, oriented posteriad; surface rather uneven and covered with moderately large shallow punctures which are mostly irregularly distributed, and with short, semierect setae which are oriented posteriad. Leg (Fig. 5 F) slender; coxa elongate; femur robust, elongate, flattened, obliquely attached to trochanter, distinctly wider than tibia, with short cavity for reception of tibia; tibia elongate, very slightly longer than femur, apically with two short spurs; tarsus slender, distinctly shorter than tibia in fore legs, less distinctly shorter in mid legs, and only slightly shorter in hind legs; tarsomeres 1–4 progressively shorter, tarsomere 1 longest, longer than combined lengths of tarsomeres 2 and 3, tarsomere 4 shortest, medially emarginate, ventrally with membrane, tarsomere 5 elongate, narrower than preceding tarsomeres; pretarsal claws simple, slightly curved. Abdomen (Fig. 4 B) with all ventrites densely covered with fine punctures and moderately long, semi-erect, posteriorly directed setae; tergites IX and X (Fig. 6 A, B) tightly connected by membrane, tergite IX slightly more than twice as wide as long, anteromedially deeply and narrowly emarginate to receive base of tergite X, densely covered with moderately long setae; tergite X weakly sclerotized, ~ 1.4 × as wide as long, apicomedially slightly produced; sternite IX (Fig. 6 C) elongate, ~ 1.8 × as long as wide, gradually slightly widened toward apex, widest subapically, apically subtruncate and medially slightly emarginate. Male genitalia (Fig. 6 D – F) ~ 2 × as long as maximum phallobase width; median lobe ~ 1.5 × as long as phallobase, not surpassing apices of parameres, elongate, slender, widest basally, subparallel-sided for most of its length and then narrowed at apical 3 / 5 of their length, narrowest subapically, apically rounded, basal struts short and distinctly divergent; parameres elongate, robust, apically membranous and narrowly rounded; phallobase transverse, ~ 1.3 × as wide as long, U-shaped. Females and immature stages unknown. Distribution. Philippines.
Language: English
Domain: Life Sciences
Open Access status: green
OpenAlex ID: W7225770761
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