Poa longifolia Trin., s. s.
Year: 2026
Type: article
Abstract: 1.Poa longifoliaTrin. s. s., Mém. Acad. Imp. Sci. Saint-Pétersbourg, Sér. 6, Sci. Math., Seconde Pt. Sci. Nat. 4, 2 (1): 61–62. 1836. Type protologue V. spp. Cauc. Type specimen. 14 Jul 1829; Caucasus: In locis graminoisis vs. montem Pagun, 3,000 m, C. A. Meyer s. n. (lectotype, designated in Tzvelev 1976, p. 463: (LE [ex hb. TRIN, 433 - c 2, rjs IMG _ 1906. jpg]; isolectotype: LE [TRIN microfiche 433 - c 3, K photo]. syntype (?): K (photo of LE plant C. A. Meyer no. 90, 14 Jul 1829 –1830., image!), LE [TRIN 433 - c 3 “ P. sudetica β ” “ C. A. Meyer no. 90 ”, K photo, image!]). Description. Perennials; hermaphroditic; tufted, without short rhizomes, usually dense, green; tillers numerous, all or mainly intravaginal; bulbs absent. Culms 50–100 (– 120) cm tall, erect, relatively robust, smooth or very lightly scabrous below nodes, with 1–2 (– 3) nodes exposed, lower nodes 1–2.3 mm diam. Leaves basal and cauline; leaf sheaths compressed, keel without a wing, lightly scabrous, glabrous, basal sheaths persistent; flag-leaf sheaths 7–15 cm long, margins fused 60 % the length; ligules 1–2 mm long, scarious, watery-white, abaxially smooth or scabrous, upper margin distinctly asperous, apices truncate to slightly rounded, sometimes peaked in the middle; ligules of sterile shoots ca. 0.5 mm, not noticeably decurrent; blades of upper culm 8–15 cm long, (1.2 –) 2–3 mm wide, graduated distally, flat or folded, moderately thick, persistent, abaxially smooth or lightly scabrous along the keel, adaxially lightly to moderately scabrous mainly costally, margins scabrous, apices prow-tipped; flag-leaf blades as long or longer than their sheaths; sterile shoot blades to 40 cm long, narrow, folded to involute, 1.2–3 mm wide, fairly firm. Panicles (4 –) 7–13 (– 17) cm long, erect or slightly lax, contracted or loosely contracted, fairly congested (with over 50 spikelets), proximal internode 2.5–3.5 cm long; branches 2–5 at the lowest nodes; primary branches 2–5 cm long, with 4–10 spikelets per branch, erect to ascending, moderately slender, weakly to distinctly angled, moderately to densely scabrous mainly on the angles or all over; pedicels mostly less than 1 / 2 their spikelets in length. Spikelets 5–7 (– 8) mm long, lanceolate, strongly compressed, not bulbiferous; florets 2–4 (– 5); rachilla internodes ca. 0.8–1.2 (– 1.5) mm long, glabrous, smooth or scaberulous; glumes unequal to subequal, slightly thinner and shinier than lemmas, keels and veins scabrous, surfaces distally scabrous, margins scabrous, apices pointed; lower glumes 2.3–3.5 mm long, narrowly lanceolate, (1 –) 3 - veined; upper glumes 3–4 mm long, lanceolate, 3 - veined; callus glabrous (infrequently with a brief, sparse dorsal web on proximal floret); lemmas (3.5 –) 4.2–4.5 (– 5.5) mm long, 5 - veined, lanceolate, 5 - veined, distally violaceous, strongly keeled, usually glabrous, keels and marginal veins scabrous, keels of proximal lemma (s) sometimes short villous in the proximal 1 / 8 th, intercostal surfaces distally lightly scaberulous, intermediate veins distinct to prominent, margins narrowly hyaline, edges distinctly scaberulous, apices acute, pointed; palea keels scabrous in distal 2 / 3, spinules short (0.02–0.05 mm), dense, inter-keel area muriculate to sparsely scaberulous. Flowers bisexual; lodicules 0.5–0.6 mm long, broadly lanceolate, with a narrow lateral lobe, glabrous; anthers 2.5–3.5 mm long. Caryopses 1.3 mm long, pale, laterally compressed, sulcate, free from the palea; hilum 0.4 mm long ovate. 2 n = 42, 43. Clade code: Php. Distribution and ecology. Endemic to the Caucasus, specifically the Russian Caucasus and adjacent northern Transcaucasia in Georgia and Azerbaijan (see Grossheim, 1939, Karta 222). Northern Azerbaijan, Georgia, Russia. Caucasus and northern Transcaucasia. Forested and high open steppe and meadows to alpine. Conservation status. None. Comments. Edmondson (1985) treated Poa longifolia in a broad sense, including P. controversa, P. meyeri, P. paryadrica as synonyms. Poa longifolia subspecies longifolia has not been verified from Turkey. A syntype, LE ex LE TRIN [LE TRIN 433 - c 1], Rv. Baidara, 15 Sep {1929}, C. A. Meyer, was annotated as the Type by E. Gabrieljan in 1957, but see Tzvelev, loc. cit.) [rjs LE IMG _ 1905. jpg]. Original material of P. longifolia (LE ex LE TRIN [LE TRIN 433 - c 1; rjs IMG _ 1905. jpg]) “ In graminosis ad rivulorum Baidara, (1000 ….), 15 Sep, C. A. Meyer ”, was annotated as Typus by E. Gabrieljan in 1957, and proposed by “ Gabrieljan (l. c. 71) ” {This is an error for Gabrieljan 1961, p. 72} according to Tzvelev (1976, p. 463). Baidara River is in the Georgian Caucasus, the specimen was annotated as Poa iberica by Litvinow. Tzvelev rejected that typification as not matching the protologue. Regarding Armenian plants cited by Gabrieljan 2010, some or all are Poa paryadrica. Tzvelev selected the specimen labeled by Trinius as “ Poa longifolia m. ” In RJS’s opinion Poa longifolia s. s. is misapplied in Turkey and to at least some material in Turkey and Armenia that should now be considered to be Poa paryadrica, and other material as Poa longifolia subsp. meyeri. Poa iberica is reported as diploid (Edmondson 1975 as “ longifolia ”, but the Stavropol garden source plant confirmed as P. iberica by Soreng et al. 7900), and tetraploid (Tzvelev and Probatova 2019), and P. longifolia as hexaploid (Tzvelev and Probatova 2019), while neither name, P. meyeri nor P. longifolia subsp. meyeri, was accounted for in Russia. Gabrieljan (2010) accepted both as species for Armenia. RJS considered both options and here adopted subspecies ranks as some plants were judged to be intermediate even in the Russan Caucasus. Grossheim (1939) considered P. longifolia to be restricted to the Caucasus (Carta No. 222), and mapped specimens from Caucasus and south through Transcaucasia including Armenian Turkey as P. meyeri (Carta No. 118) and P. iberica (Carta No. 223). All three were reported for Turkey (Edmondson 1985, as P. iberica, P. longifolia subsp. longifolia and meyeri), but he doubted P. iberica occurred there. We agree with Grossheim that P. longifolia does not occur in Turkey, and consider material of “ Poa iberica ” in Turkey, and at least some of the Transcaucasian material called that, to belong to P. paryadrica. Representative specimens. Russia, Caucasus (Soreng et al. 7945 - a *; 7946; 7957).
Language: English
Domain: Life Sciences
Open Access status: green
OpenAlex ID: W7225813007
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